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Flanker Justin Tipuric will stunt Wales as head coach Warren Gatland makes 13 adjustments due to their final World Cup pool match against Uruguay.
Centre Hadleigh Parkes and wing Josh Adams will be the only players remaining from Wednesdays win over Fiji.
Lock Adam Beard makes his very first World Cup appearance.
Wales have achieved quarter-final eligibility and need just two points to finish high in Pool D.
Wales were always planning to make changes for this fixture, provided the turnaround from Wednesdays bonus-point victory against Fiji.
The need was compounded with the amount of injuries picked up together with facility Jonathan Davies and Dan Biggar forced off the field at Oita, during that game.
Gatland said George North took an ankle knock in the Fiji game and the need to rest Adams was donated to by again starting despite the latter suffering a leg in that game.
Adams switches from the wing together Hallam Amos producing his very first tournament appearance.
Scrum-half Aled Davies, hooker Ryan Elias and lock Bradley Davies additionally earn their first taste of activity in Japan.
Spinning and injuries mean Wales have chosen Tomos Williams and Gareth Davies as their backs that are only on the bench, along with six forward.
That means there isnt going to be a specialist fly-half cover Rhys Patchell, that begins at 10.
Gatland has indicated the cover will probably come in Parkes, who has Amos, who has never played 10 within his mature career, or limited experience of playing professional rugby in fly-half.
The replacement scrum-halves may be requested to pay for the position.
The Wales coach states having four days involving the Fiji and Uruguay matches has enforceda bit of a juggling act and selection discussions comprised asking flanker James Davies to perform at the backs.
We knew that right from the beginning we were planning to expose a few gamers, said Gatland.
I spoke before the World Cup [to say] we would probably go into this match with only one 10 anyway so we are searching for cover there also.
It has been tough with the short turnaround, but weve got a forward pack that are all fresh and fairly excited and they are excited about the opportunity.
The message to the players for the match against Uruguay is the door isnt closed. There are opportunities for players to venture out there and impress and stake a claim for the quarter-finals.
Gatland added:We spoke about if we might have put James Davies from the backs. He has lots of sevens expertise and Justin (Tipuric) can be fairly comfortable playing in the wider channels also.
Were convinced we have cover with players and having the ability to move around if we need to do that.
Weve had two gamers back upward in Hadleigh Parkes and Josh Adams.
Ideally we would have liked to have given Josh a rest, but has not been the case and thats the reason why weve ended up opting for a 6-2 divide with 2 nines covering the springs.
Gareth (Davies) went onto the wing at the end of the Fiji game and did a great job to us. We now have two caliber nines who can cover different positions.
Wales: Halfpenny; Adams, Watkin, Parkes, Amos; Patchell, A Davies; Smith, Elias, Lewis, B Davies, Beard, Shingler, Tipuric (capt), Wainwright.
Replacements: G Davies, Dee, Carre, W Jones, Ball, Moriarty T Williams.
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